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She is letting the other dogs know that she is alpha bitch in that group. If the other dog thinks otherwise then there will be a stand off and weak one will back off. It's just her temperament, the owner needs to be aware of this and not let it get out of control.
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Interesting, Storm, the bitch in question playfights more with Ella and humps her, but runs off when Zee instigates play with her. Ella is a more nervous temperament than Zee. And if Storm and Ella are having a spat, Zee will intervene and tell off the one doing the bossing, like a peace-maker. Does that mean Zee is alpha?